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Anther developmental periods were classified as previously described for Arabidopsis ( 2.8 ROS detection The Nitroblue tetrazolium chloride (NBT) and 3,3’-diaminobenzidine (DAB) staining were performed according to a published method ( 1 O 2 was measured using Singlet Oxygen Sensor Green reagent (Beijing Biolab Technology Co., Ltd., Beijing, China) according to the manufacturer’s instructions